The development of a benthic ecosystem on a sandy bottom at 3m depth in the saline Lake Grevelingen, is described. The development started after the closure of the former Grevelingen estuary and was followed for 4.5 yrs. Ecosystem attributes studied were biomass, secondary production, production/biomass ratio, type of food chains, species diversity, pattern diversity, size of organisms, type of life-cycles, and r-and K-selection. The results are discussed in relation to E. P. Odum's (1969) theory of ecosystem development.
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